Allow `autosave: true` to be used with inverse of

With the changes in #25337, double save bugs are pretty much impossible,
so we can just lift this restriction with pretty much no change. There
were a handful of cases where we were relying on specific quirks in
tests that had to be updated. The change to has_one associations was due
to a particularly interesting test where an autosaved has_one
association was replaced with a new child, where the child failed to
save but the test wanted to check that the parent id persisted to `nil`.

I think this is almost certainly the wrong behavior, and I may change
that behavior later. But ultimately the root cause was because we never
remove the parent in memory when nullifying the child. This makes #23197
no longer needed, but it is what we'll do to fix some issues on 5.0

Close #23197

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